The carpel is homologous with the megasporophylls of certain spore-bearing vascular plants (pteridophytes) and the ovuliferous scales of gymnosperms.

心皮和能产生孢子的维管植（类）还有具胚珠的子植的大孢子源的。

Bract scale The structure found in large numbers in the female cone of gymnosperms, each bearing an ovuliferous scale in its axil.

在子植的雌球果中大量存在的结构，每个苞鳞都包含一个珠鳞。