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Spedding and his colleagues think that the unique aerodynamic wake signatures of bat wings are caused by different mechanical operations in the upstroke of the wingbeat.

斯佩丁和他同事们认为，独特后，蝙蝠翅膀所造成不同机械操作，在upstrokewingbeat 。

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原声例句
TED-Ed（视频版）

Hummingbirds however, turn their wrists during the upstroke, changing their wing's angle to flap in a figure 8 pattern that continually generates lift throughout their wingbeats.

然而，上击时转动手腕，改变翅膀的角度，以 8 字形模式拍打，整个拍翅程中不断产生升力。

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PBS地球-动物趣闻

Their wing joints are also stiffer than those of flying birds, which helps them produce thrust on both the downstroke - like most birds do - and on the upstroke of the wing.

它们的翅膀也比飞的硬，这有助于它们像大多数类一样下击和上击时产生推力。

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