One inclusion formed somewhat later exterior to the exosporium. Another inclusion was formed within the exosporium and remained with the spore after mother cell lysis.

当芽胞成细胞裂解后，一种晶体被释放出来，成自由的伴胞晶体，另一种在芽胞外壁内侧成，成熟后被包裹在芽胞外壁以内不脱落，始终和芽胞粘连在一起。