pugmark

Paw print, or pugmark, tracing is another common technique for counting tigers.

爪印和足迹样是计算数量方法。

The researchers then use the pictures to measure the length and width of each pugmark, the angle between toes, and 10 other variables to establish an ID.

研究人员这些照片来测量脚印长度和宽度以及脚趾之间角度，以及10种其他变量来确定不“身份”。